śabdaḥ — sound; sparśaḥ — touch; rasaḥ — taste; gandhaḥ — fragrance; rūpam — form; ca — and; iti — thus; artha — of sense objects; jātayaḥ — the categories; gati — movement; ukti — speech; utsarga — excretion (by both the genitals and anus); śilpāni — and manufacture; karma-āyatana — by the above-mentioned working senses; siddhayaḥ — accomplished.