putrān — to his sons; adhyāpayat — he taught; tān — those Vedas; tu — and; brahma-ṛṣīn — to the great sages among the brāhmaṇas; brahma — in the art of Vedic recitation; kovidān — who were very expert; te — they; tu — moreover; dharma — in religious rituals; upadeṣṭāraḥ — instructors; sva-putrebhyaḥ — to their own sons; samādiśan — imparted.